Contains 4 accepted taxa overall.
Characteristics
The genera Botrypus (B. virginianus) and Sceptridium (S. biternatum, S. jenmanii, S. lunarioides) are now commonly segregated from Botrychium (Shinohara et al. 2013; Dauphin et al. 2014; PPG I 2016).
Classification
Citation
BOTRYCHIUM Swartz, J. Bot. (Schrader) 1800(2): 8, 110. 1802.
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TYPE: Botrychium lunaria (Linnaeus) Swartz (Osmunda lunaria Linnaeus) Lectotypified by Smith, Hist. Filicum 369. 1875.
Species
| Scientific Name | Common Name | Herbarium Specimens | Status | Photos |
|---|---|---|---|---|
| Botrychium biternatum | SOUTHERN GRAPE-FERN | Native FAC (NWPL) | ||
| Botrychium jenmanii | ALABAMA GRAPE-FERN; JENMAN'S GRAPE-FERN | Native FAC (NWPL) | ||
| Botrychium lunarioides | WINTER GRAPE-FERN | Native | ||
| Botrychium virginianum | RATTLESNAKE FERN | Native FACU (NWPL) |
[Family identification key]
1. Petiole of fertile leaf long-exserted from the ground, the sterile green branches subsessile and attached at the base of the rachis of the fertile blades; leaves generally finely dissected with the pinnules mostly lanceolate; leaves present mostly Mar-Oct (Botrypus)
1. Petiole of fertile leaf short, the rachis of the sterile green blade long and forking distant from where the base of the rachis of the fertile blade; leaves moderately dissected with the pinnules mostly suborbicular, elliptic, or rounded; leaves present year-round or present mostly Oct-Apr (Sceptridium)
2
2. Roots pale or yellowish, up 30 main roots; leaves prostrate or nearly so, present mostly Oct-Apr, commonly 2 per plant; sporophore stalks and midrib broadly flattened, fleshy; leaves
2. Roots dark or brownish, up to 15 main roots; leaves erect to ascending, present year-round, commonly 1-2 per plants; sporophore stalks and midrib only slightly flattened, not fleshy
3
3. Basal pinnae mostly subopposite; pinnule venation pinnate, with strong midrib; blades bluish green or green to dark green
3. Basal pinnae remotely alternate; pinnule venation nearly like ribs of fan but with short midrib; blades dull gray-green
Genus Synonyms
| Synonym | Full Citation | Basionym | Type |
|---|---|---|---|
| Botrypus | Botrypus Michaux, Fl. Bor.-Amer. 2: 274. 1803. | TYPE: Botrypus virginicus Michaux, nom.. illegit. (Osmunda virginica Linnaeus, nom. illegit.; Osmunda virginiana Linnaeus) Lectotypified by Pfeiffer, Nomencl. 1: 449. 1873; Pichi Sermolli, Webbia 26: 497. 1972. | |
| Japanobotrychum | Japanobotrychum Masamune, J. Soc. Trop. Agric. 3: 246. 1931. | TYPE: Japanobotrychum arisanense Masamune | |
| Osmundopteris | Osmundopteris (Milde) Small, Ferns. S.E. States 377, 482. 1938. | BASIONYM: Botrychium Linnaeus, sect. Osmundopteris Milde, Filices Eur. 209. 1867. | TYPE: Osmundopteris virginiana (Linnaeus) Small (Osmunda virginiana Linnaeus) = Botrypus Michaux 1803, by lectotypification. |
| Sceptridium | Sceptridium Lyon, Bot. Gaz. 40: 457. 1905. | TYPE: Sceptridium obliquum (Muhlenberg ex Willdenow) Lyon (Botrychium obliquum Muhlenberg ex Willdenow) Lectotypified by R. T. Clausen, Mem. Torrey Bot. Club 19(2): 24. 1938. |